Maternal t | Embryo t + 1 | Endosperm t + 1 | ||||
---|---|---|---|---|---|---|

Genotype | Symbol | Values | Genotype | Symbol | Value | |

QQ(t) | QQ(t + 1) | 1 | μ_{1} = μ + 2a | QQQ(t + 1) | 1 | μ_{1} = μ + |

Qq(t) | QQ(t + 1) | 2 | μ_{2} = μ + a + β_{1} | QQQ(t + 1) | 2 | μ_{2} = μ + + β_{1} |

Qq(t + 1) | 3 | μ_{3} = μ + β_{2} | QQq(t + 1) | 3 | μ_{3} = μ + + β_{2} | |

qQ(t + 1) | 3 | μ_{3} = μ + β_{2} | Qqq(t + 1) | 4 | μ_{4} = μ − + β_{3} | |

qq(t + 1) | 4 | μ_{4} = μ − a + β_{3} | qqq(t + 1) | 5 | μ_{5} = μ − + β_{4} | |

qq(t) | qq(t + 1) | 5 | μ_{5} = μ − 2a | qqq(t + 1) | 6 | μ_{6} = μ − |